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Uneven crumb -- proofing or other problem?

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I made these loaves based on the San Joaquin recipe by dmsnyder, but I upped the whole grain content to 50% (mostly wheat, but also a significant amount of rye), and raised the hydration (by an unmeasured amount, but it ended up feeling higher than I had intended). I only cut one of the loaves so far, but as you can see, the crumb was pretty irregular. Is this a sign of a particular part of the process I can improve on? Proofing? Something else? Or is it just normal variation? It seems a little too drastic for that last option.

Spelt Whole Wheat Rice Ricotta Bread

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Since I started my new job in the big city I have not had a lot of time to bake.  Last weekend I found enough time to whip this one together.  I used mostly freshed milled and sifted flours for this bake and added a little KAF Bread flour for some extra strength.  I have grown to love adding cooked rice to the bread.  You don't taste the actual rice but it adds a great texture to the crumb and helps promote softness as well.  The ricotta cheese also helps create a soft and moist crumb which this one certainly had.

Tinned H5GL with AYW

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I decided to bake Hamelman's 5 Grain Levain in a tin and mix in some AYW.  This was in an attempt to get more oven spring, and prevent sagging sides.  I used 20% AYW of total water, and shaped (somewhat unsuccessfully) for a tin.  The result was quite pleasing, but I suspect the tin resulted in a slightly gummy crumb, so next time I will remove the loaf and bake free-form for the final 10 minutes.  I also used baking spray on the loaf top to prevent the plastic bag from sticking during retard.  This resulted in a very dark top crus

1-2-3 Third Attempt

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Decided on upping the starter to 150 so...

Starter 150g

Water 300

Flour 225 AP/225 BF

12g of salt 

Mixed it all up, let it rest for an hour

Looked at it and folded it every 2 hours for 6.5 hours 

Wonderful marshmallow texture when I put it in the bowl

Let it go in the fridge for 22 hours

Baked at the usual 450 for 500, I did forget to dunk the parchment paper in water. 

123 the III

3-Way Oat Sourdough

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This bread was so good, I thought it was worthy of creating my first blog!  The original recipe came from Rose Levy Berenbaum's site, here; she cites original recipe from Chef Andrew Meltzer.  The version she posts is a yeasted loaf and was spectacularly good.  I made it as she posted and below is the result.  The dough contains an oat porridge, toasted oats, and the loaves are rolled in oats before baking.  I was obviously a little more successful in rolling one

Whole Grain Sourdough Bread. 3

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Week three and bread three. Last time, i felt the structure of the dough was lacking, so I did a few more stretch and folds before shaping. To facilitate fermentation, I also did some more folds early and slightly decreased the salt. In this i was certainly right, for the first time, the dough doubled in bulk overnight and proofed very well.